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重新安装浏览器，或使用别的浏览器Drosophila Smt3 negatively regulates JNK signaling through sequestering Hipk in the nucleus.
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):2477-85. doi: 10.1242/dev.061770. Epub
2011 May 11.Drosophila Smt3 negatively regulates JNK signaling through sequestering Hipk in the nucleus.1, , , , , , , , .1State Key Laboratory of Brain and Cognitive Science, Institute of Biophysics, Chinese Academy of Sciences, Datun Road 15, Beijing 100101, China.AbstractPost-translational modification by the small ubiquitin-related modifier (SUMO) is important for a variety of cellular and developmental processes. However, the precise mechanism(s) that connects sumoylation to specific developmental signaling pathways remains relatively less clear. Here, we show that Smt3 knockdown in Drosophila wing discs causes phenotypes resembling JNK gain of function, including ectopic apoptosis and apoptosis-induced compensatory growth. Smt3 depletion leads to an increased expression of JNK target genes Mmp1 and puckered. We show that, although knockdown of the homeodomain-interacting protein kinase (Hipk) suppresses Smt3 depletion-induced activation of JNK, Hipk overexpression synergistically enhances this type of JNK activation. We further demonstrate that Hipk is sumolylated in vivo, and its nuclear localization is dependent on the sumoylation pathway. Our results thus establish a mechanistic connection between the sumoylation pathway and the JNK pathway through the action of Hipk. We propose that the sumoylation-controlled balance between cytoplasmic and nuclear Hipk plays a crucial role in regulating JNK signaling.PMID:
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bi卑鄙beibian北边beibu北部,被捕beican悲惨beicha备查beidaihe北戴河beidao被盗beidao'erchi背道而驰beidaxiyang北大西洋beidili背地里beidong被动beifang北方beifen备份,悲愤beifeng北风beifu背负,被俘beigan倍感beigao被告beigaoren被告人beigeng备耕beigongsheyi杯弓蛇影beiguan悲观beiguo北国beihai北海,被害beihao备好beihou背后beihuang备荒beiji北极beijia倍加beijian备件beijiang北疆beijie被劫beijing北京,背景beijingdaxue北京大学beijinglixia背井离乡beijingren北京人beijingshi北京市beijingshiji北京时间beijingzhan北京站beijixing北极星beiju悲剧beike备课beikong被控beikou被扣beilei蓓蕾beili背离beiliao备料beilie卑劣beimei北美beimian背面,北面beipan背叛beipo被迫beiqi备齐beiru被褥beisha被杀beishang悲伤beishou倍受beishu倍数beishuichexi杯水车薪beishuiyizha背水一战beitong悲痛beiwanglu备忘录beiwei北纬beixia卑下beixinqiyi背信弃义beiyin背阴beiyong备用beiyongjin备用金beiyou备有beiyue北约beiyuwei被誉为beizeng倍增beizhan备战beizhu备注beizhuang悲壮ben本,苯,奔,笨,贲,畚,坌,锛,夯,体benbao本报benbo奔波benbu本部benbumen本部门benchang本厂benchi奔驰benci本次bendanwei本单位bendi本地bendiqu本地区benfang奔放benfu奔赴beng泵,崩,绷,甭,蹦,甏,迸,嘣,蚌bengai本该bengbu蚌埠bengfa迸发bengkui崩溃benglie崩裂benglong崩龙bengta崩塌benguo本国benhang本行benjie本届benjin本金benkan本刊benke本科benkesheng本科生benlai本来benlaimianmu本来面目benli本利benling本领benliu奔流benmang奔忙benmodaozhi本末倒置benneng本能benniandu本年度benpao奔跑benqian本钱benqu本区benren本人bensang奔丧bense本色benshai本色benshen本身bensheng本省benshi本事,本市benshiji本世纪benshijichu本世纪初benshijimo本世纪末benshijinei本世纪内benteng奔腾bentou奔头bentu本土benwei本位benwen本文benxian本县benxiang奔向benxiao本校benxing本性benxitong本系统benyi本意benying本应benyue本月benzhe本着benzhi本质,本职benzhigongzu本职工作benzhong笨重benzhou本周benzhuo笨拙benzou奔走benzouhuhao奔走呼号benzouxiangg奔走相告bi比,必,壁,笔,臂,闭,逼,彼,避,毕,弊,鼻,碧,币,辟,吡,蔽,铋,俾,鄙,哔,毙,痹,璧,庇,匕,婢,敝,秕,篦,箅,陛,弼,荜,筚,畀,狴,裨,荸,嬖,滗,舭,襞,髀,泌,秘,濞,薜,萆,毖,拂,跸,庳,愎,妣,蓖,贲,埤,睥,芘,稗,蚍bi'an彼岸bi'erbutan避而不谈bian便,边,变,编,遍,辨,辩,扁,鞭,贬,辫,匾,煸,弁,汴,卞,缏,鳊,砭,苄,忭,笾,碥,蝙,褊,窆bianbenjiali变本加厉bianbie辨别bianbiezhenj辨别真假bianbo辩驳bianbu遍布biance鞭策biancheng变成,编成bianchui边陲biancuan编篡biandang便当biandao编导biande变得biandi遍地,贬低biandiansuo变电所biandianzhan变电站biandong变动biandui编队bianfang边防bianfeiweiba变废为宝bianganbianx边干边学biange变革bianguan边关bianhan便函bianhao编号,变好bianhei变黑bianhu辩护bianhua变化bianhuaduodu变化多端bianhuai变坏bianhuamoce变化莫测bianhuan变换,变幻bianhuanmoce变幻莫测bianhuanwuch变幻无常bianhuawucha变化无常bianhulvshi辩护律师bianhuren辩护人bianji编辑,遍及,边际bianjiang边疆bianjibu编辑部bianjichuban编辑出版bianjie边界bianjiexian边界线bianjing边境bianli便利bianlun辩论bianma编码bianmin便民bianneng便能biannuan变暖bianpai编排bianqing辨清bianqu边区bianren辨认bianru编入bianse变色biantai变态biantong变通bianwai编外bianwei变为bianweihui编委会bianxian边线bianxiang变相bianxie编写bianxieshi便携式bianxing变形bianyan边沿bianyang变样bianyaqi变压器bianye遍野bianyi编译,贬义,便宜bianyin编印bianyu便于,编余bianyuan边缘,边远bianzao编造bianzhe编者bianzhe'an编者按bianzheng辩证bianzhengbia边整边改bianzhengfa辩证法bianzhi编制,编织,贬值,变质,贬职bianzhong变种bianzhu编著bianzhuan编撰bianzi辫子bianzu编组biao表,标,彪,鳔,膘,裱,飙,婊,镖,杓,飚,髟,飑,瘭,镳,骠,罴,嘌biaobai表白biaobang标榜biaoben标本biaobing标兵biaoceng表层biaoda表达biaoge表格biaoji标记biaojia标价biaojue表决biaojuequan表决权biaolibuyi表里不一biaoliruyi表里如一biaolu表露biaomian表面biaomianhua表面化biaoming表明,标明biaopi表皮biaoqian标签biaoqing表情biaoshi表示,标示,标识biaoshuai表率biaotai表态biaoti标题biaoxian表现biaoxiang表象biaoxianli表现力biaoxinliyi标新立异biaoyan表演biaoyang表扬biaoyu标语biaoyupai标语牌biaozhang表彰biaozhanghui表彰会biaozhi标志,标致,标识biaozhun标准bibai必败bibei必备bibijieshi比比皆是bibing弊病bibo鄙薄,碧波bibukemian必不可免bibukeshao必不可少bici彼此bicizhijian彼此之间bida笔答bide必得bidei必得bideshang比得上bidiao笔调biding必定biduan弊端biduolishao弊多利少bie别,憋,瘪,蹩,鳖biechu别处biechuxincai别出心裁bieguo别国biejujiangxi别具匠心biejutese别具特色biejuyige别具一格biekaishengm别开生面biekan别看bieli别离bieniu别扭bieren别人bieshuyizhi别树一帜bieti别提biewuchulu别无出路biewusuoqiu别无所求biewutafa别无他法biewuxuanze别无选择bieyang别样bieyoufengwe别有风味bieyouyongxi别有用心biezhi别致bifa笔法bifang比方bifen比分bifeng避风bifenggang避风港biganzi笔杆子bigongbijing毕恭毕敬biguansuoguo闭关锁国biguanzishou闭关自守bihe闭合bihu庇护bihua壁画,比划bihui闭会biji笔记bijia比价bijiang必将bijiao比较bijiao'eryan比较而言bijiben笔记本bijie毕节bijin逼近bijing毕竟bijingzhidi必经之地bikai避开bikong碧空bilan碧蓝bilei壁垒bili比例bilu笔录bilv比率,碧绿bimengeng闭门羹bimenzaoche闭门造车bimian避免bimo笔墨bimu闭幕bimuci闭幕词bimushi闭幕式bin宾,滨,斌,彬,鬓,濒,摈,殡,豳,膑,缤,玢,傧,镔,槟,髌binan避难binansuo避难所binbinyouli彬彬有礼binfen缤纷bing并,病,兵,丙,冰,炳,柄,饼,禀,秉,摒,屏,邴,疒,槟bing'an病案,并按bingbao冰雹,禀报,并报bingbian病变,兵变bingbu并不bingbudengyu并不等于bingbumaodun并不矛盾bingbuneng并不能bingbushi并不是bingcan病残bingcheng秉承bingchonghai病虫害bingchuan冰川bingcongkour病从口入bingcun并存bingdong冰冻bingdu病毒bingfa并发bingfei并非bingfeiruci并非如此bingfeiyishi并非易事binggan饼干binggen病根binggong秉公binggong'erd秉公而断binggongbans秉公办事binggongchan兵工厂binggui冰柜bingguishens兵贵神速binghe冰河bingjian并肩bingjiaqiqu并驾齐驱bingjin并进bingju并举bingjun病菌bingleng冰冷bingli兵力,病例,并立,病历,病理binglian并联bingliang冰凉binglie并列bingma兵马bingmeiyou并没有bingmo病魔bingneng并能bingpai并排bingqi摒弃,兵器bingqiangmaz兵强马壮bingqie并且bingqing并请bingren病人bingru并入bingta病榻bingtai病态bingti并提bingtianxued冰天雪地bingtong病痛bingtuan兵团bingtun并吞binguan宾馆bingwangfadi并网发电bingwei病危,并未bingwu并无bingxiang冰箱bingxing并行,禀性,秉性bingxue冰雪bingyao并要bingyi兵役bingyifa兵役法bingyin病因bingying兵营bingyong并用bingyou并有bingyu病愈bingyuan兵员,病员,病院bingzhan兵站bingzheng病症bingzhi秉直bingzhong兵种bingzhuan并转bingzhuang病状bini比拟binke宾客binlin濒临binqi摈弃binwei濒危binyu濒于binzhirugui宾至如归binzhu宾主bipo逼迫biran必然biranxing必然性biru比如birushuo比如说bisai比赛bise闭塞bishangbuzu比上不足bishangliang逼上梁山bisheng毕生,必胜bishi鄙视,笔试,比试bishou匕首bishu避暑bishushanzhu避暑山庄biting笔挺biwu笔误,比武bixia笔下bixiayouyu比下有余bixiu必修bixiuke必修课bixu必须,必需bixupin必需品biyao必要biyaoxing必要性biye毕业biyeban毕业班biyesheng毕业生biyi笔译biyou必有biyouzhilu必由之路biyu比喻,碧玉biyun避孕biyunyao避孕药bizhao比照bizhe笔者,比着bizhen逼真bizhi笔直bizhong比重bizhongjiuqi避重就轻bizhu比诸bo波,薄,伯,播,拨,剥,博,泊,铂,驳,玻,勃,脖,舶,帛,搏,钵,簸,箔,跛,啵,饽,渤,膊,钹,擘,卜,踣,菠,礴,亳,魄,鹁,檗,孛,掰,趵,般,柏,白,薜,荸,坡bo'ai博爱bo'erdala博尔塔拉bobo勃勃bocaizhongch博采众长bochi驳斥bochu播出bochuan驳船bodajingshen博大精深bode博得bodong拨动bodou搏斗boduan波段boduo剥夺bofa播发bofang播放bogei拨给bogutongjin博古通今bohai渤海bohui驳回boji搏击bojiang播讲bokuan拨款bolan波澜,博览bolang波浪bolanhui博览会boli剥离,薄利,玻璃boliduoxiao薄利多销bolin柏林boluanfanzhe拨乱反正boluo剥落bonong拨弄borandanu勃然大怒borong拨冗boruo薄弱boshi博士boshisheng博士生bosong播送botao波涛botong拨通bowei泊位bowu博物bowuguan博物馆bowuyuan博物院boxue剥削,博学boxueduocai博学多才boyin波音,播音boying播映boyinyuan播音员bozhe波折bozhong播种bu不,部,布,步,补,捕,卜,簿,钚,埠,钸,怖,哺,卟,堡,逋,醭,晡,瓿,埔bu'an不安bubai不败bubaizhidi不败之地buban部颁bubanbiaozhu部颁标准bubao补报bubaofenpei不包分配bubeibukang不卑不亢bubi不必bubian不便,不变bubing步兵bubingshi步兵师bubuweiying步步为营buceng不曾buchang补偿,不畅buchangfei补偿费buchengbili不成比例buchengtiton不成体统buchenzhi不称职buchixiawen不耻下问buchong补充buchusuoliao不出所料buci'erbie不辞而别bucixinku不辞辛苦bucuo不错buda不大budakeneng不大可能budan不但,不单budang不当budangzhichu不当之处budaode不道德budaozhichu不到之处bude'erzhi不得而知budeliao不得了budeng不等budengyu不等于buderenxin不得人心budeyaoling不得要领budeyi不得已budiao步调budiaoyizhi步调一致buding不定budingqi不定期budiyu不低于budong不懂budongchan不动产buduan不断budui部队,不对buduojian不多见bufa步伐,不乏,补发bufada不发达bufafenzi不法分子bufan不凡bufang不妨bufangyishi不妨一试bufaqiren不乏其人bufaxingwei不法行为bufazhitu不法之徒bufen部分bufengzhuoyi捕风捉影bufenshengfu不分胜负bufu不符,不服bufucunzai不复存在bufuqi不服气bufushu不服输bufuzeren不负责任bufuzhongwan不负众望bugai不该bugan不敢buganbujing不干不净bugandang不敢当buganjimo不甘寂寞buganluohou不甘落后buganshiruo不甘示弱buganxin不甘心bugao布告bugao'erci不告而辞bugong不公bugou不够bugu不顾buguan不管buguang不光bugui不轨buguize不规则buguo不过buguoguan不过关buguyiqie不顾一切buhao不好buhaoyisi不好意思buhe不和,不合buhefa不合法buheli不合理buheshiyi不合时宜buhuaihaoyi不怀好意buhuan'ersan不欢而散buhuangbuman不慌不忙buhuo捕获buhuoliang捕获量buhuozhinian不惑之年buji不及,补给,不计,部级bujia不佳bujiafenxi不加分析bujian部件,不见bujiandan不简单bujiandangni不减当年bujiang不讲bujianjingzh不见经传bujiao补交bujiaobuzao不骄不躁bujiasisuo不加思索bujibaochou不计报酬bujie不解bujiezhiyuan不解之缘bujiguan部机关bujin不仅,不禁,不尽bujinbuman不紧不慢bujing不经bujing'erzou不胫而走bujingji不经济bujingqi不景气bujingyishi不经一事bujinjin不仅仅bujinran不尽然bujinrenqing不近人情bujinrenyi不尽人意bujinruci不仅如此bujiqishu不计其数bujitian不几天bujiu不久,补救bujiuqian不久前bujiyu不急于buju布局,不拘bujuan不倦bujue不觉,不绝bujueyu'er不绝于耳bujuyige不拘一格bukan不堪bukangbubei不亢不卑bukanshexian不堪设想bukao补考,不靠buke补课,不可bukebu不可不bukeduode不可多得bukegaoren不可告人bukekaijiao不可开交buken不肯bukeneng不可能bukeqi不客气bukexue不科学bukeyishi不可一世bukuai不快bukui不愧bulaiwang不来往bulang布朗bulao捕捞bulao'erhuo不劳而获bulei部类buli不利,不力buliang不良buliao不料buliaojie不了解buliaoliaozh不了了之buliyu不利于bulun不论bulunbulei不伦不类buluo部落bulv步履bulvweijian步履维艰buman不满,布满bumang不忙bumaozhidi不毛之地bumen部门bumian不免bumiao不妙buming不明bumingbubai不明不白bumou'erhe不谋而合bumousili不谋私利bunan不难buneng不能bunengbu不能不bunengziba不能自拔bupa不怕bupei不配bupianbuyi不偏不倚bupin补品buping不平bupingdeng不平等bupingheng不平衡buqi'eryu不期而遇buqiang步枪buqiao不巧buqieshiji不切实际buqiushangji不求上进buqiushenjie不求甚解buqu不屈buqubunao不屈不挠buque补缺buran不然burandehua不然的话buren不忍burong不容burongxu不容许buru不如,步入,哺乳,补入busha捕杀bushan不善bushangbuxia不上不下bushao不少bushen不慎,不深busheng不胜bushengmeiju不胜枚举bushi不适,不时,捕食,不失bushidati不识大体bushishiji不失时机bushishiwu不识时务bushiwei不失为bushiying不适应bushiyongwul不使用武力bushizhici不实之词bushou部首bushouhuanyi不受欢迎bushu部署,部属,不熟bushufu不舒服bushui补税bushuyu不属于busuan不算busuzhike不速之客butie补贴butiefei补贴费butong不同,不通butongfanxia不同凡响butongyi不同意butongyu不同于butongzhichu不同之处butuo不妥buwai不外buwaihu不外乎buwang不忘buwanquanton不完全统计buwei部委,部位,不畏buwen不稳,不问buwending不稳定buwudaoli不无道理buwuguanxi不无关系buwuzhengye不务正业buxi不惜,补习,不息buxia部下buxiang不详,不想,不象buxiangchen不相称buxianggan不相干buxiangrong不相容buxiangshang不相上下buxiangshiyi不相适应buxiangzhizh不祥之兆buxie不懈,布鞋,补写buxieyigu不屑一顾buxigongben不惜工本buxin不信buxing不行,不幸,步行buxingle不行了buxiu不朽buxu不许,不需buxuan补选buxue补血buxuewushu不学无术buxunsiqing不徇私情buyan不严buyan'eryu不言而喻buyang哺养buyanqifan不厌其烦buyanziming不言自明buyao不要buyaojin不要紧buyayu不亚于buyi不易,不宜,补益,不一,不已,不依,不意,补遗,不义,布依buyi'erfei不翼而飞buyiding不一定buying不应buyinggai不应该buyiweiran不以为然buyiyuli不遗余力buyizhicai不义之财buyong不用buyongshuo不用说buyou部优,不由buyoude不由得buyoufenshuo不由分说buyouzizhu不由自主buyu哺育,不予,捕鱼buyuan不愿,不远buyuanqianli不远千里buyue不悦buyue'ertong不约而同buyukaolv不予考虑buyukuai不愉快buzai不再,不在buzaicilie不在此列buzaihu不在乎buzaihuaxia不在话下buzaobuwan不早不晚buzeshouduan不择手段buzhang部长buzhangji部长级buzhebukou不折不扣buzhen不振buzheng不正buzhengzhife不正之风buzhi不止,布置,不只,不值,不致,不知buzhibujue不知不觉buzhidao不知道buzhide不值得buzhikefou不置可否buzhiquxiang不知去向buzhisuocuo不知所措buzhiyici不止一次buzhiyiti不值一提buzhiyu不至于buzhizhiju不智之举buzhizhizhen不治之症buzhou步骤,不周buzhu补助buzhufei补助费buzhujin补助金buzhun不准buzhuo捕捉buzi步子buziliangli不自量力buzouguochan不走过场buzu不足,补足,部族buzuguachi不足挂齿buzuweiping不足为凭buzuweiqi不足为奇buzuzhichu不足之处ca擦,嚓,礤,拆,搽cai才,采,菜,财,材,彩,裁,猜,踩,蔡,睬caibian采编caice猜测caichan财产caichang菜场caichanquan财产权caideng彩灯caidi菜地caidian彩电caiding裁定caifa采伐caifang采访caifu财富caigan才干caigou采购caigouyuan采购员caihong彩虹caihua才华caihuachuzho才华出众caihuahengyi才华横溢caiji采集caijia菜价caijian裁减caijing财经caijue裁决,采掘caijun裁军caikuai财会caikuang采矿caikuangye采矿业cailanzi菜篮子caili财力cailiao材料cailiaochang材料厂cailiaofe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